Guardian deities/Manga

Pokémon Adventures
The guardian deities first appeared in the. They were first seen in a fantasy in The Delivery of Rotom and the Girl, where Hala, Olivia, and Nanu each discussed that all four deities were spotted by people on their respective island. As the guardian deities were normally reclusive, the Kahunas assumed that something must have angered the four Pokémon. In The Final Match and a Surprising Finale!, was given the task of delivering a special Mirage Berry to each guardian deity as a way of calming them down.

Tapu Koko first appeared in The Delivery of Rotom and the Girl, attacking Sun and. After a short battle, it escaped, leaving behind a Sparkling Stone for Sun. In The Decision and the Tournament of Six, Tapu Koko appeared at Mahalo Trail, where it saved Moon and Nebby after the latter broke the Plank Bridge to escape from a group of that attacked it.

Tapu Lele first appeared in True Identity and the Totem Pokémon of Brooklet Hill, where it fought against a that tried to escape from an Ultra Wormhole and lost. It was saved by Moon, who fired a -tipped arrow at Xurkitree, forcing it to retreat back to where it came from. Afterward, it returned to the Ruins of Life to recuperate, only for Gladion to arrive and try to capture it. After being defeated by Sun, Gladion gave up on Tapu Lele, claiming that it was too weak to fight against the Ultra Beasts. Once Gladion left, Sun successfully delivered the Mirage Berry to Tapu Lele. As thanks, Tapu Lele decided to test the abilities of Sun and his friends by battling them. Though it proved to be powerful, Tapu Lele left satisfied after Sun managed to land a hit on it with his newly-evolved,.

Tapu Bulu first appeared at Po Town, where it rescued, , and from an attacking. Sun was initially reluctant to give a Mirage Berry to Tapu Bulu, as he felt that it had no reason to help humans as one was responsible for the appearance of Ultra Beasts in Alola to begin with. Moon argued that if humans were the cause, then they should take responsibility by fixing the mess as well. Sun and Moon then give the Mirage Berry to Tapu Bulu, which it readily accepts.

Tapu Fini first appeared at Seafolk Village, where it watched Hapu battle a group of wild led by a. Tapu Fini watched the battle to see if Hapu was worthy of succeeding the title of Kahuna of Poni Island. After Hapu successfully defeated the wild Pokémon, Sun had her deliver the Mirage Berry, as he was too injured by Crabominable's attacks to do so himself. Hapu then received a Sparklng Stone from Tapu Fini, confirming that it accepted her as the next Kahuna.